Can PCIe x4 fit in a PCIe x16 slot?

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According to PCIE Expresscard , a x1 card can be inserted into the x4, x8, and x16 connectors 15 This is referred to as up PCIe x4 devices are compatible with larger PCIe slots such as x8 and x16, and can be installed in these slots as well However, the bandwidth of the device is

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